Lactococcus lactis (MG1363)

Food-grade, GRAS lactic acid bacterium used for centuries in dairy fermentation. Non-colonizing and ideal for ingestible / clinical biosensors where a food-safe, gut-transient host is wanted.

BSL-1GRASprobiotic

Identity

Species
Lactococcus lactis
Strain
MG1363
NCBI taxid
416870
Genome
NC_009004.1

Safety

Biosafety level
BSL-1
Pathogenic
No
GRAS
Yes — FDA GRAS for food use; EFSA QPS; centuries of safe use in dairy.
Probiotic
Yes — Food fermentation; engineered L. lactis trialed clinically (e.g. mucosal delivery).
Biocontainment
thyA-based biological containment demonstrated in clinical trials (transient survival).
BSL-1 basis
ABSA Risk Group 1; FDA GRAS; EFSA QPS.

Gram-positive, no LPS endotoxin; transient gut passage (does not colonize).

Traits

Gram
positive
Oxygen
facultative
Optimal temp
30 °C
Doubling time
~60 min
Spore-forming
No
Transformable
Yes

Engineering

CRISPR tooling
dCas9/CRISPRi demonstrated in lactic acid bacteria.
Common promoters
PnisA (nisin-inducible, NICE system), P32 (constitutive)
Selection markers
ErmR, CmR
Toolkits
NICE (nisin-controlled expression), SEVA-compatible vectors

Biosensors using this chassis

Fluoride sensor (riboswitch + CRISPRa)

Detects fluoride in drinking water using a natural fluoride riboswitch as the sensor, gating a CRISPRa circuit driving a fluorescent reporter, in the food-grade GRAS chassis Lactococcus lactis.

environmentalCRISPRa-activation

Gut acid/pH sensor (GadE + CRISPRa)

Reports gut acidity / low pH (linked to short-chain-fatty-acid load and dysbiosis). The acid-responsive Gad system gates a CRISPRa circuit driving a fluorescent reporter, in the food-grade acid-tolerant chassis Lactococcus lactis.

clinical-gutCRISPRa-activation

Lactate sensor (LldR + CRISPRi)

Detects L-lactate, a fermentation and clinical metabolite. The LldR lactate-responsive regulator gates a CRISPRa circuit driving a fluorescent reporter, in the food-grade GRAS chassis Lactococcus lactis.

chemicalCRISPRa-activation

Naringenin/flavonoid sensor (FdeR + CRISPRa)

Detects naringenin and related flavonoids (dietary and nutraceutical compounds). The Marionette FdeR sensor gates a CRISPRa circuit driving a fluorescent reporter, in the food-grade GRAS chassis Lactococcus lactis.

chemicalCRISPRa-activation

Streptococcus sensor (CSP/ComD + CRISPRa)

Detects Streptococcus via its competence-stimulating peptide (CSP). The ComD/ComE quorum system gates a CRISPRa circuit driving a fluorescent reporter, in the food-grade Gram-positive chassis Lactococcus lactis.

pathogenCRISPRa-activation

Tryptophan sensor (TrpR + CRISPRi)

Detects tryptophan, a dietary amino acid and precursor of gut-microbial indole/serotonin metabolites. The classic TrpR repressor gates a CRISPRi circuit driving a fluorescent reporter, in the food-grade GRAS chassis Lactococcus lactis.

clinical-gutCRISPRi-repression

Zinc sensor (ZitR + CRISPRi)

Detects zinc, a nutritional and water-quality marker. The native L. lactis ZitR zinc regulator gates a CRISPRi circuit driving a fluorescent reporter, in the food-grade GRAS chassis Lactococcus lactis.

chemicalCRISPRi-repression