Tetracycline sensor (TetR + CRISPRi)

Detects tetracycline antibiotic contamination in water and food. The classic TetR sensor gates a CRISPRi circuit driving a visible pigment, in safe E. coli K-12.

Chemical / metaboliteBSL-1 chassistemplatetetracyclineantibioticantibiotic-pollutionwaterfood-safetychemicalenvironmentalCRISPRi
Input
Tetracycline / anhydrotetracycline (aTc)
Chemical / metabolite
Sense
CRISPRi-repression
dCas9 (S. pyogenes, catalytically dead)
Chassis
E. coli K-12 (MG1655)
BSL-1
Output
amilCP
pigment

What it detects

Analyte
Tetracycline / anhydrotetracycline (aTc) — TetR responds from low nM aTc
Category
Chemical / metabolite
Signal
Tetracycline-class antibiotic residues in water, food, and wastewater

Genetic circuit

⤢ click to enlarge

Genetic construct (SBOL)

The DNA construct as transcription units, drawn with SBOL Visual part glyphs.

⤢ click to enlarge

CRISPR sensing mechanism

Strategy
CRISPRi-repression · NOT logic
Cas protein
dCas9 (S. pyogenes, catalytically dead)
Analyte sensor
TetR de-represses the Ptet promoter on binding tetracycline/aTc.
Signal flow
Tetracycline -> TetR releases Ptet -> transcribes an anti-pigment sgRNA -> CRISPRi represses a constitutive amilCP cassette -> pigment fades with antibiotic (NOT). Pair an inverter for colour-on.

Safe chassis

E. coli K-12 (MG1655)Escherichia coli

The non-pathogenic laboratory workhorse. K-12 strains have lost the ability to colonize the human gut and are the reference BSL-1 host for genetic engineering, with the deepest tooling of any bacterial chassis.

BSL-1

Genetic parts

PartRoleSource / id
TetR regulator
Tetracycline/aTc-responsive repressor; one of the best-characterised regulators.
regulatorTn10 tet operon (also a Marionette/standard sensor)
Ptet promoter
De-repressed by tetracycline-bound TetR.
promoterBBa_R0040
Anti-pigment sgRNAsgRNAdesigned against the amilCP promoter
sgRNA scaffold (SpCas9)
GTTTTAGAGCTAGAAATAGCAAGTTAAAATAAGGCTAGTCCGTTATCAACTTGAAAAAGTGGCACCGAGTCGGTGC
sgRNAStandard SpCas9 scaffold
dCas9dCas9Qi et al. 2013 (CRISPRi)
amilCP chromoproteinreporterBBa_K592009

Output & readout

Type
pigment
Reporter
amilCP
Readout
Visible pigment (naked eye / smartphone)
Positive result
Pigment change reports tetracycline contamination.

Performance

Limit of detection
TetR module: low-nM aTc (module-validated).
Dynamic range
~1-1000 nM tetracycline/aTc
Response time
~150 min
Device validated
No — design template (parts validated individually)

TetR/Ptet is among the most characterised regulator-promoter pairs in biology; integrated CRISPRi device is a design template.

Safety

Biosafety level
BSL-1 (non-pathogenic chassis)
GRAS chassis
No
Biocontainment
Lab/contained; add kill-switch for field use.
Field-deployable
Yes (with containment)

E. coli K-12 non-pathogenic BSL-1 host; relevant to antibiotic-pollution surveillance. Note: do not select the host with tetracycline resistance.

Build & run

#StageStep
1designDesign anti-pigment sgRNA
Target the amilCP promoter; check host off-targets.
2assemblyAssemble units
TU1: TetR + Ptet -> sgRNA. TU2: dCas9 + constitutive amilCP. Avoid TetR-resistance markers.
3transformationTransform E. coli K-12
Select (non-tet marker); confirm baseline pigment without antibiotic.
4inductionExpose to sample
Add water/food sample + tetracycline standard curve.
5readoutScore colour
Compare pigment to the curve.

Source & parts

Design
Design template combining the classic TetR tetracycline sensor with E. coli CRISPRi
Parts validated in
  • Hillen & Berens / TetR tetracycline regulation (Tn10)
  • Qi et al. 2013, Cell (CRISPRi)
License
Parts per their original sources; design template CC BY 4.0