Gut inflammation memory recorder (tetrathionate)

Records a transient gut-inflammation episode and remembers it. A recombinase memory switch, triggered by the tetrathionate sensor, flips a DNA state that persists after the signal is gone — so a single exposure is still detectable days later in stool.

Clinical / gut biomarkermemory circuitBSL-1 chassistemplateclinical-gutinflammationtetrathionatememoryrecorderrecombinaseprobioticNissle
Input
Tetrathionate (transient)
Clinical / gut biomarker
Sense
CRISPRi-repression
dCas9 (S. pyogenes, catalytically dead)
Chassis
E. coli Nissle 1917
BSL-1
Output
sfGFP
fluorescent

What it detects

memory design — records a transient exposure and remembers it.

Analyte
Tetrathionate (transient) — Records exposure above the sensor's threshold
Category
Clinical / gut biomarker
Signal
A transient tetrathionate / inflammation episode that may not coincide with sampling

Genetic circuit

⤢ click to enlarge

Genetic construct (SBOL)

The DNA construct as transcription units, drawn with SBOL Visual part glyphs.

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CRISPR sensing mechanism

Strategy
CRISPRi-repression · BUFFER logic
Cas protein
dCas9 (S. pyogenes, catalytically dead)
Analyte sensor
The TtrS/TtrR tetrathionate sensor drives a recombinase (integrase) that flips a DNA register; the flipped state is heritable and stable without further signal.
Signal flow
Tetrathionate -> TtrS/TtrR expresses a recombinase -> the recombinase inverts a DNA segment (set the memory bit) -> the flipped state switches a reporter permanently ON. CRISPRi can gate or reset the register. The record persists after the signal clears.

Safe chassis

E. coli Nissle 1917Escherichia coli

A probiotic E. coli used in humans for over a century (Mutaflor). Colonizes the gut safely, making it the chassis of choice for clinical / gut biomarker biosensors.

BSL-1probiotic

Genetic parts

PartRoleSource / id
TtrS/TtrR sensor
Validated in vivo; triggers the recombinase.
regulatorDaeffler/Riglar et al. (tetrathionate sensor in Nissle)
Recombinase / integrase (e.g. Bxb1)
Flips a DNA register to store the memory bit.
cdsphage serine integrase
Flippable DNA register
Heritable two-state memory element.
operatorattB/attP-flanked invertible segment
dCas9
Optional gating/reset of the register.
dCas9Qi et al. 2013 (CRISPRi)
sgRNA scaffold (SpCas9)
GTTTTAGAGCTAGAAATAGCAAGTTAAAATAAGGCTAGTCCGTTATCAACTTGAAAAAGTGGCACCGAGTCGGTGC
sgRNAStandard SpCas9 scaffold
sfGFP reporter
Switched permanently ON by the flipped register; recoverable from stool.
reporterPedelacq et al. 2006

Output & readout

Type
fluorescent
Reporter
sfGFP
Readout
Fluorescence (flow cytometry on recovered cells)
Positive result
Cells stay fluorescent after a past inflammation episode, even once tetrathionate is gone.

Performance

Device validated
No — design template (parts validated individually)

Recombinase memory + the tetrathionate sensor are each validated in vivo (Riglar et al. 2017); this CRISPRi-gated memory variant is a design template. Captures intermittent inflammation that real-time sensors miss.

Safety

Biosafety level
BSL-1 (non-pathogenic chassis)
GRAS chassis
No
Biocontainment
Probiotic E. coli Nissle host; add thyA/dapA auxotrophy for gut-restricted containment.
Field-deployable
Lab / supervised use

Probiotic chassis with a human-safety record; research / supervised clinical use only.

Build & run

#StageStep
1designDesign the memory register
Place a recombinase-invertible register controlling the reporter; design any CRISPRi reset.
2assemblyAssemble units
TU1: TtrS/TtrR -> recombinase. TU2: invertible register -> reporter. Optional dCas9 reset arm.
3transformationTransform E. coli Nissle 1917
Select; verify the register flips on tetrathionate and stays flipped.
4inductionPulse exposure
Apply a transient tetrathionate pulse, then remove it; confirm the memory persists.
5readoutRecover and read state
Recover cells (e.g. from stool) and read the flipped fraction.

Source & parts

Design
Design template: a recombinase memory recorder triggered by the validated tetrathionate sensor, with optional CRISPRi gating
Parts validated in
  • Riglar et al. 2017, Nat. Biotechnol. (in-vivo memory sensor)
  • Bxb1 / serine integrase memory devices
  • Qi et al. 2013, Cell (CRISPRi)
License
Parts per their original sources; design template CC BY 4.0