Estrogen / endocrine-disruptor sensor (ERα + CRISPRa)

Detects estrogens and estrogenic endocrine-disrupting chemicals in water. Uses the human estrogen receptor (ERα) expressed in GRAS baker's yeast — a eukaryotic sensing class bacteria cannot do — gating a CRISPRa circuit driving a fluorescent reporter.

Chemical / metaboliteBSL-1 chassisGRAStemplateestrogenendocrine-disruptorEDCwaterchemicaleukaryoticyeastCRISPRa
Input
Estrogens / estrogenic EDCs
Chemical / metabolite
Sense
CRISPRa-activation
dCas9-VPR (eukaryotic CRISPRa activator)
Chassis
Saccharomyces cerevisiae (BY4741)
BSL-1
Output
yEGFP
fluorescent

What it detects

Analyte
Estrogens / estrogenic EDCs — Yeast ERα assays detect sub-nM to nM 17β-estradiol
Category
Chemical / metabolite
Signal
17β-estradiol and estrogen-mimicking endocrine disruptors in water

Genetic circuit

⤢ click to enlarge

Genetic construct (SBOL)

The DNA construct as transcription units, drawn with SBOL Visual part glyphs.

⤢ click to enlarge

CRISPR sensing mechanism

Strategy
CRISPRa-activation · amplifier logic
Cas protein
dCas9-VPR (eukaryotic CRISPRa activator)
Analyte sensor
Ligand-bound human ERα binds estrogen-response elements (EREs), activating an ERE-controlled promoter (the classic yeast estrogen screen).
Signal flow
Estrogen -> ERα binds ERE promoter -> transcribes an sgRNA -> dCas9-VPR amplifies a fluorescent reporter (CRISPRa) -> fluorescence rises with estrogenic activity.

Safe chassis

Saccharomyces cerevisiae (BY4741)Saccharomyces cerevisiae

GRAS baker's yeast — the model eukaryote. Being a eukaryote, it can host human nuclear receptors and GPCRs, unlocking sensing of steroids, hormones, and other eukaryotic-specific ligands that bacteria cannot detect. dCas9 CRISPRi/CRISPRa are well established.

BSL-1GRAS · FDA GRAS

Genetic parts

PartRoleSource / id
Human estrogen receptor alpha (ERα)
Ligand-activated nuclear receptor — requires a eukaryotic host.
regulatorHuman ESR1, expressed in yeast (Routledge & Sumpter 1996, yeast estrogen screen)
ERE promoter
Activated by ligand-bound ERα.
promoterEstrogen-response-element-controlled yeast promoter
Reporter-activating sgRNAsgRNAdesigned for CRISPRa upstream of a weak yeast promoter
sgRNA scaffold (SpCas9)
GTTTTAGAGCTAGAAATAGCAAGTTAAAATAAGGCTAGTCCGTTATCAACTTGAAAAAGTGGCACCGAGTCGGTGC
sgRNAStandard SpCas9 scaffold
dCas9-VPR
Eukaryotic transcriptional activator.
dCas9Yeast CRISPRa (dCas9 fused to VPR/VP64)
yEGFPreporteryeast-enhanced GFP

Output & readout

Type
fluorescent
Reporter
yEGFP
Readout
Green fluorescence (plate reader)
Positive result
Fluorescence increases with estrogenic activity.

Performance

Limit of detection
Yeast ERα assays: sub-nM 17β-estradiol (module-validated).
Dynamic range
~0.1-10 nM estradiol equivalents
Response time
~360 min
Device validated
No — design template (parts validated individually)

The yeast estrogen screen (ERα + ERE) is a validated assay; the CRISPRa-amplified version here is a design template. Reports total estrogenic activity (a key advantage for unknown EDC mixtures).

Safety

Biosafety level
BSL-1 (non-pathogenic chassis)
GRAS chassis
Yes
Biocontainment
Auxotrophic markers; GRAS yeast host.
Field-deployable
Yes (with containment)

GRAS baker's yeast; eukaryotic chassis enables human-receptor-based sensing.

Build & run

#StageStep
1designDesign CRISPRa sgRNA
Target a weak yeast promoter; check S. cerevisiae off-targets.
2assemblyAssemble units
Express ERα; ERE -> sgRNA; dCas9-VPR; weak promoter -> yEGFP. Use the Yeast Toolkit (MoClo).
3transformationTransform S. cerevisiae
Integrate at defined loci; select on auxotrophic markers.
4inductionExpose to sample
Incubate with the water sample + estradiol standard curve.
5readoutMeasure fluorescence
Report estradiol-equivalent estrogenic activity.

Source & parts

Design
Design template combining the yeast estrogen screen (ERα/ERE) with eukaryotic CRISPRa
Parts validated in
  • Routledge & Sumpter 1996 (yeast estrogen screen, ERα/ERE)
  • Yeast dCas9-VPR CRISPRa literature
License
Parts per their original sources; design template CC BY 4.0